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Mechanics of limb bone loading during terrestrial locomotion in river cooter turtles (Pseudemys concinna). Butcher MT, Blob RW. 2008. Journal of Experimental Biology, 211: 1187-1202.

Myosin isoform fiber type and fiber size in the tail of the Virginia opossum (Didelphis virginiana). Hazimihalis PJ*, Gorvet MA*, Butcher MT. 2013. Anatomical Record, 296: 96-107.

Architectural specializations of the intrinsic thoracic limb musculature of the American badger (Taxidea taxus). Moore AL*, Budny JE*, Russell AP, Butcher MT. 2013. Journal of Morphology, 274: 35-48.   

*indicates student author from my lab

Femoral loading mechanics in Virginia opossums (Didelphis virginiana): torsion and mediolateral bending in mammalian parasagittal locomotion. Gosnell WC, Butcher MT, Maie T, Blob RW. 2011. Journal of Experimental Biology, 214: 3455-3466.

In vivo strain in the femur of the opossum (Didelphis virginiana) during terrestrial locomotion: testing hypotheses of evolutionary shifts in bone loading and design. Butcher MT, Hudzik NB*, White BJ*, Gosnell WC, Parrish JHA, Blob RW. 2011. Journal of Experimental Biology, 214: 2631-2640.

Locomotor loading mechanics in the hindlimbs of tegu lizards (Tupinambis merianae): comparative      and evolutionary implications. Sheffield KM, Butcher MT, Shugart SK, Gander JC, Blob RW. 2011. Journal of Experimental Biology, 214: 2616-2630.

Contractile properties of muscle fibers from the forelimb deep and superficial digital flexors of horses. Butcher MT, Chase PB, Hermanson JW, Clark AN, Brunet NM, Bertram JEA. 2010. American Journal of Physiology, (Regulatory Integrative Comparative Physiology), 299: R996-R1005.

Contractile behavior of the forelimb digital flexors during steady-state locomotion in horses (Equus caballus): an initial test of muscle architectural hypotheses about in vivo function. Butcher MT, Hermanson JW, Ducharme NG, Mitchell LM, Soderholm LV, Bertram JEA. 2009. Comparative Biochemistry Physiology A, Molecular & Integrative Physiology, 152: 100-114.

Correlation of bone loading and muscle contractile function in the hindlimb of the river cooter turtle (Pseudemys concinna). Aiello BR*, Blob RW, Butcher MT. 2013. Journal of Morphology, 274: 1060-1069.

Development of an integrative biomechanics course for STEM majors. Panta YM, Butcher MT. 2012. Proceedings of the American Society for Engineering Education. Download: PDF

In vivo strains in the femur of river cooter turtles (Pseudemys concinna) during terrestrial locomotion: tests of force-platform models of loading mechanics. Butcher MT, Espinoza NR, Cirilo SR, Blob RW. 2008. Journal of Experimental Biology, 211: 2397-2407.

Most papers are available for free PDF download at PubMed







Step-length and required friction walking. Cooper RC, Butcher MT, Bertram JEA. 2008. Gait Posture, 27: 547-551.

Superficial digital flexor tendon lesions in racehorses as a sequelae to muscle fatigue: a preliminary study. Butcher MT, Hermanson JW, Ducharme NG, Mitchell LM, Soderholm LV, Bertram JEA. 2007. Equine Veterinary Journal, 39: 540-545.


Collateral ligaments of the distal sesamoid bone in the digit of Equus: Re-evaluating mid-stance function. Butcher MT, Bertram JEA, Bezuidenhout AJ. 2006. Journal of Morphology, 267: 1177-1185.

Constrained optimization in human running. Gutmann AK, Butcher MT, Jacobi B, Bertram JEA. 2006. Journal of Experimental Biology, 209: 622-632.


Jump distance increases while carrying hand-held weights: Impulse, history and jump performance in a simple lab exercise. Butcher MT, Bertram JEA. 2004. Journal of Science Education and Technology, 13: 285-297. Download: PDF


Age differences in fetlock joint kinematics of Thoroughbreds. Butcher MT, Ashley-Ross MA. 2002. Journal of Biomechanics, 35: 563-571.


Muscle architecture and out-force potential of the thoracic limb in the Eastern mole (Scalopus aquaticus). Rose JA*, Sandefur M, Huskey S, Demler J*, Butcher MT. Journal of Morphology, 274: 1277-1287.

Collision based dynamics of human walking and running in relation to other mammalian gaits. Lee DV, Tutor NC, Butcher MT, Bertram JEA. 2013. Proceedings of the Royal Society B, 280: 20131779.

Myosin heavy chain (MHC) isoform expression in the prehensile tails of didelphid marsupials: functional differences between arboreal and terrestrial opossums. Rupert JE*, Moriera A, Cordero Schmidt  E, VandeBerg JL, Rodriguez Herrera B, Butcher MT. Anatomical Record, accepted 01-12-2014.

Functional osteological of the forelimb digging apparatus of badgers. Rose JA*, Moore AL*, Russell AP, Butcher MT. Journal of Mammalogy, in revision.